第 15 节
作者:猜火车      更新:2021-02-24 23:33      字数:9322
  with their respiration which is effected by the skin alone。
  A worm after swallowing earth; whether for making its burrow or for
  food; soon comes to the surface to empty its body。  The ejected
  earth is thoroughly mingled with the intestinal secretions; and is
  thus rendered viscid。  After being dried it sets hard。  I have
  watched worms during the act of ejection; and when the earth was in
  a very liquid state it was ejected in little spurts; and by a slow
  peristaltic movement when not so liquid。  It is not cast
  indifferently on any side; but with some care; first on one and
  then on another side; the tail being used almost like a trowel。
  When a worm comes to the surface to eject earth; the tail
  protrudes; but when it collects leaves its head must protrude。
  Worms therefore must have the power of turning round in their
  closely…fitting burrows; and this; as it appears to us; would be a
  difficult feat。  As soon as a little heap has been formed; the worm
  apparently avoids; for the sake of safety; protruding its tail; and
  the earthy matter is forced up through the previously deposited
  soft mass。  The mouth of the same burrow is used for this purpose
  for a considerable time。  In the case of the tower…like castings
  (see Fig。 2) near Nice; and of the similar but still taller towers
  from Bengal (hereafter to be described and figured); a considerable
  degree of skill is exhibited in their construction。  Dr。 King also
  observed that the passage up these towers hardly ever ran in the
  same exact line with the underlying burrow; so that a thin
  cylindrical object such as a haulm of grass; could not be passed
  down the tower into the burrow; and this change of direction
  probably serves in some manner as a protection。
  Worms do not always eject their castings on the surface of the
  ground。  When they can find any cavity; as when burrowing in newly
  turned…up earth; or between the stems of banked…up plants; they
  deposit their castings in such places。  So again any hollow beneath
  a large stone lying on the surface of the ground; is soon filled up
  with their castings。  According to Hensen; old burrows are
  habitually used for this purpose; but as far as my experience
  serves; this is not the case; excepting with those near the surface
  in recently dug ground。  I think that Hensen may have been deceived
  by the walls of old burrows; lined with black earth; having sunk in
  or collapsed; for black streaks are thus left; and these are
  conspicuous when passing through light…coloured soil; and might be
  mistaken for completely filled…up burrows。
  It is certain that old burrows collapse in the course of time; for
  as we shall see in the next chapter; the fine earth voided by
  worms; if spread out uniformly; would form in many places in the
  course of a year a layer 0。2 of an inch in thickness; so that at
  any rate this large amount is not deposited within the old unused
  burrows。  If the burrows did not collapse; the whole ground would
  be first thickly riddled with holes to a depth of about ten inches;
  and in fifty years a hollow unsupported space; ten inches in depth;
  would be left。  The holes left by the decay of successively formed
  roots of trees and plants must likewise collapse in the course of
  time。
  The burrows of worms run down perpendicularly or a little
  obliquely; and where the soil is at all argillaceous; there is no
  difficulty in believing that the walls would slowly flow or slide
  inwards during very wet weather。  When; however; the soil is sandy
  or mingled with many small stones; it can hardly be viscous enough
  to flow inwards during even the wettest weather; but another agency
  may here come into play。  After much rain the ground swells; and as
  it cannot expand laterally; the surface rises; during dry weather
  it sinks again。  For instance; a large flat stone laid on the
  surface of a field sank 3。33 mm。 whilst the weather was dry between
  May 9th and June 13th; and rose 1。91 mm; between September 7th and
  19th of the same year; much rain having fallen during the latter
  part of this time。  During frosts and thaws the movements were
  twice as great。  These observations were made by my son Horace; who
  will hereafter publish an account of the movements of this stone
  during successive wet and dry seasons; and of the effects of its
  being undermined by worms。  Now when the ground swells; if it be
  penetrated by cylindrical holes; such as worm…burrows; their walls
  will tend to yield and be pressed inwards; and the yielding will be
  greater in the deeper parts (supposing the whole to be equally
  moistened) from the greater weight of the superincumbent soil which
  has to be raised; than in the parts near the surface。  When the
  ground dries; the walls will shrink a little and the burrows will
  be a little enlarged。  Their enlargement; however; through the
  lateral contraction of the ground; will not be favoured; but rather
  opposed; by the weight of the superincumbent soil。
  Distribution of Worms。Earth…worms are found in all parts of the
  world; and some of the genera have an enormous range。 {41}  They
  inhabit the most isolated islands; they abound in Iceland; and are
  known to exist in the West Indies; St。 Helena; Madagascar; New
  Caledonia and Tahiti。  In the Antarctic regions; worms from
  Kerguelen Land have been described by Ray Lankester; and I found
  them in the Falkland Islands。  How they reach such isolated islands
  is at present quite unknown。  They are easily killed by salt…water;
  and it does not appear probable that young worms or their egg…
  capsules could be carried in earth adhering to the feet or beaks of
  land…birds。  Moreover Kerguelen Land is not now inhabited by any
  land…bird。
  In this volume we are chiefly concerned with the earth cast up by
  worms; and I have gleaned a few facts on this subject with respect
  to distant lands。  Worms throw up plenty of castings in the United
  States。  In Venezuela; castings; probably ejected by species of
  Urochaeta; are common in the gardens and fields; but not in the
  forests; as I hear from Dr。 Ernst of Caracas。  He collected 156
  castings from the court…yard of his house; having an area of 200
  square yards。  They varied in bulk from half a cubic centimeter to
  five cubic centimeters; and were on an average three cubic
  centimeters。  They were; therefore; of small size in comparison
  with those often found in England; for six large castings from a
  field near my house averaged 16 cubic centimeters。  Several species
  of earth…worms are common in St。 Catharina in South Brazil; and
  Fritz Muller informs me 〃that in most parts of the forests and
  pasture…lands; the whole soil; to a depth of a quarter of a metre;
  looks as if it had passed repeatedly through the intestines of
  earth…worms; even where hardly any castings are to be seen on the
  surface。〃  A gigantic but very rare species is found there; the
  burrows of which are sometimes even two centimeters or nearly 0。8
  of an inch in diameter; and which apparently penetrate the ground
  to a great depth。
  In the dry climate of New South Wales; I hardly expected that worms
  would be common; but Dr。 G。 Krefft of Sydney; to whom I applied;
  after making inquiries from gardeners and others; and from his own
  observations; informs me that their castings abound。  He sent me
  some collected after heavy rain; and they consisted of little
  pellets; about 0。15 inch in diameter; and the blackened sandy earth
  of which they were formed still cohered with considerable tenacity。
  The late Mr。 John Scott of the Botanic Gardens near Calcutta made
  many observations for me on worms living under the hot and humid
  climate of Bengal。  The castings abound almost everywhere; in
  jungles and in the open ground; to a greater degree; as he thinks;
  than in England。  After the water has subsided from the flooded
  rice…fields; the whole surface very soon becomes studded with
  castingsa fact which much surprised Mr。 Scott; as he did not know
  how long worms could survive beneath water。  They cause much
  trouble in the Botanic garden; 〃for some of the finest of our lawns
  can be kept in anything like order only by being almost daily
  rolled; if left undisturbed for a few days they become studded with
  large castings。〃  These closely resemble those described as
  abounding near Nice; and they are probably the work of a species of
  Perichaeta。  They stand up like towers; with an open passage in the
  centre。
  A figure of one of these castings from a photograph is here given
  (Fig。 3)。  The largest received by me was 3。5 inches in height and
  1。35 inch in diameter; another was only 0。75 inch in diameter and
  2。75 in height。  In the following year; Mr。 Scott measured several
  of the largest; one was 6 inches in height and nearly 1。5 in
  diameter:  two others were 5 inches in height and respectively 2
  and rather more than 2。5 inches in diameter。  The average weight of
  the 22 castings sent to me was 35 grammes (1。25 oz。); and one of
  them weighed 44。8 grammes (or 2 oz。)。 All these castings were
  thrown up either in one night or in two。  Where the ground in
  Bengal is dry; as under large trees; casting