第 13 节
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猜火车 更新:2021-02-24 23:33 字数:9322
forwards into this part; which consequently swells and pushes away
the earth on all sides。 The anterior extremity thus serves as a
wedge。 It also serves; as we have before seen; for prehension and
suction; and as a tactile organ。 A worm was placed on loose mould;
and it buried itself in between two and three minutes。 On another
occasion four worms disappeared in 15 minutes between the sides of
the pot and the earth; which had been moderately pressed down。 On
a third occasion three large worms and a small one were placed on
loose mould well mixed with fine sand and firmly pressed down; and
they all disappeared; except the tail of one; in 35 minutes。 On a
fourth occasion six large worms were placed on argillaceous mud
mixed with sand firmly pressed down; and they disappeared; except
the extreme tips of the tails of two of them; in 40 minutes。 In
none of these cases; did the worms swallow; as far as could be
seen; any earth。 They generally entered the ground close to the
sides of the pot。
A pot was next filled with very fine ferruginous sand; which was
pressed down; well watered; and thus rendered extremely compact。 A
large worm left on the surface did not succeed in penetrating it
for some hours; and did not bury itself completely until 25 hrs。 40
min。 had elapsed。 This was effected by the sand being swallowed;
as was evident by the large quantity ejected from the vent; long
before the whole body had disappeared。 Castings of a similar
nature continued to be ejected from the burrow during the whole of
the following day。
As doubts have been expressed by some writers whether worms ever
swallow earth solely for the sake of making their burrows; some
additional cases may be given。 A mass of fine reddish sand; 23
inches in thickness; left on the ground for nearly two years; had
been penetrated in many places by worms; and their castings
consisted partly of the reddish sand and partly of black earth
brought up from beneath the mass。 This sand had been dug up from a
considerable depth; and was of so poor a nature that weeds could
not grow on it。 It is therefore highly improbable that it should
have been swallowed by the worms as food。 Again in a field near my
house the castings frequently consist of almost pure chalk; which
lies at only a little depth beneath the surface; and here again it
is very improbable that the chalk should have been swallowed for
the sake of the very little organic matter which could have
percolated into it from the poor overlying pasture。 Lastly; a
casting thrown up through the concrete and decayed mortar between
the tiles; with which the now ruined aisle of Beaulieu Abbey had
formerly been paved; was washed; so that the coarser matter alone
was left。 This consisted of grains of quartz; micaceous slate;
other rocks; and bricks or tiles; many of them from 1/20 to 1/10
inch in diameter。 No one will suppose that these grains were
swallowed as food; yet they formed more than half of the casting;
for they weighed 19 grains; the whole casting having weighed 33
grains。 Whenever a worm burrows to a depth of some feet in
undisturbed compact ground; it must form its passage by swallowing
the earth; for it is incredible that the ground could yield on all
sides to the pressure of the pharynx when pushed forwards within
the worm's body。
That worms swallow a larger quantity of earth for the sake of
extracting any nutritious matter which it may contain than for
making their burrows; appears to me certain。 But as this old
belief has been doubted by so high an authority as Claparede;
evidence in its favour must be given in some detail。 There is no a
priori improbability in such a belief; for besides other annelids;
especially the Arenicola marina; which throws up such a profusion
of castings on our tidal sands; and which it is believed thus
subsists; there are animals belonging to the most distinct classes;
which do not burrow; but habitually swallow large quantities of
sand; namely; the molluscan Onchidium and many Echinoderms。 {37}
If earth were swallowed only when worms deepened their burrows or
made new ones; castings would be thrown up only occasionally; but
in many places fresh castings may be seen every morning; and the
amount of earth ejected from the same burrow on successive days is
large。 Yet worms do not burrow to a great depth; except when the
weather is very dry or intensely cold。 On my lawn the black
vegetable mould or humus is only about 5 inches in thickness; and
overlies light…coloured or reddish clayey soil: now when castings
are thrown up in the greatest profusion; only a small proportion
are light coloured; and it is incredible that the worms should
daily make fresh burrows in every direction in the thin superficial
layer of dark…coloured mould; unless they obtained nutriment of
some kind from it。 I have observed a strictly analogous case in a
field near my house where bright red clay lay close beneath the
surface。 Again on one part of the Downs near Winchester the
vegetable mould overlying the chalk was found to be only from 3 to
4 inches in thickness; and the many castings here ejected were as
black as ink and did not effervesce with acids; so that the worms
must have confined themselves to this thin superficial layer of
mould; of which large quantities were daily swallowed。 In another
place at no great distance the castings were white; and why the
worms should have burrowed into the chalk in some places and not in
others; I am unable to conjecture。
Two great piles of leaves had been left to decay in my grounds; and
months after their removal; the bare surface; several yards in
diameter; was so thickly covered during several months with
castings that they formed an almost continuous layer; and the large
number of worms which lived here must have subsisted during these
months on nutritious matter contained in the black earth。
The lowest layer from another pile of decayed leaves mixed with
some earth was examined under a high power; and the number of
spores of various shapes and sizes which it contained was
astonishingly great; and these crushed in the gizzards of worms may
largely aid in supporting them。 Whenever castings are thrown up in
the greatest number; few or no leaves are drawn into the burrows;
for instance the turf along a hedgerow; about 200 yards in length;
was daily observed in the autumn during several weeks; and every
morning many fresh castings were seen; but not a single leaf was
drawn into these burrows。 These castings from their blackness and
from the nature of the subsoil could not have been brought up from
a greater depth than 6 or 8 inches。 On what could these worms have
subsisted during this whole time; if not on matter contained in the
black earth? On the other hand; whenever a large number of leaves
are drawn into the burrows; the worms seem to subsist chiefly on
them; for few earth…castings are then ejected on the surface。 This
difference in the behaviour of worms at different times; perhaps
explains a statement by Claparede; namely; that triturated leaves
and earth are always found in distinct parts of their intestines。
Worms sometimes abound in places where they can rarely or never
obtain dead or living leaves; for instance; beneath the pavement in
well…swept courtyards; into which leaves are only occasionally
blown。 My son Horace examined a house; one corner of which had
subsided; and he found here in the cellar; which was extremely
damp; many small worm…castings thrown up between the stones with
which the cellar was paved; and in this case it is improbable that
the worms could ever have obtained leaves。 Mr。 A。 C。 Horner
confirms this account; as he has seen castings in the cellars of
his house; which is an old one at Tonbridge。
But the best evidence; known to me; of worms subsisting for at
least considerable periods of time solely on the organic matter
contained in earth; is afforded by some facts communicated to me by
Dr。 King。 Near Nice large castings abound in extraordinary
numbers; so that 5 or 6 were often found within the space of a
square foot。 They consist of fine; pale…coloured earth; containing
calcareous matter; which after having passed through the bodies of
worms and being dried; coheres with considerable force。 I have
reason to believe that these castings had been formed by species of
Perichaeta; which have been naturalized here from the East。 {38}
They rise like towers; with their summits often a little broader
than their bases; sometimes to a height of above 3 and often to a
height of 2。5 inches。 The tallest of those which were measured was
3。3 inches in height and 1 inch in diameter。 A small cylindrical
passage runs up the centre of each tower; through which the worm
ascends to eject the earth which it has swallowed; and thus to add
to its height。 A structure of this kind would not allow leaves
being easily dragged from the surrounding ground into the burrows;
and Dr。 King; who looked carefully; never saw even a fragment of a
leaf thus drawn in。 N